Markus is a cruel, ruthless leader of a Katagari wolf clan. Several hundred years ago, he and a gang kidnapped and raped Bryani, who turned out to be his mate. He tried to force her to accept a mating, but she refused, so he is now impotent and very angry about it. Bryani sent their three Katagari pups (Anya, Fang, and Vane) to live with him and kept their three Arcadian children (Dare, Fury, and Star).
Several hundred years later, Acheron placed him on a deserted island with Bryani, but Vane showed up and used his powers to prevent them from killing each other.